Some more thoughts about the cipher..
In fact there is one big difference between the 340 and the 408: Although the 340 cipher text is shorter and contains more homophones, the + symbol is somehow outstanding as it appears an overall of 24 times or 7% while in the 408 the maximum count for one homophone is 16 or 4% only. Now this leads to a lot of trouble BECAUSE if we assume the + symbol not to be the only homophone for a letter, it would indicate a letter frequency of much more than 7%. Lets think about it:
Letter E: At least 5 if not 8 homophones, if sequentially used that would lead us to a letter frequency of roughly 5 times 24, thus 120 or 35%, which is unrealistic.
Letters L, T, N, O, A etc similar, assuming 3 homophones only, leading to a letter frequency of 21%.
Letters H, P etc., assuming 2 homophones only, leading to a letter frequency of - still - 14%, all of this is sort of unrealistic.
In addition to the trivial thoughts above I tried to recalculate the probability for a letter to show up three times as a double homophone. Most likely the L can be illustrated to be represented by the + symbol, however this still requires one of the homophones to represent 7% of the cipher. But we must accept that there would be two more homophones, most likely as frequent as the + symbol, so it doesn't work out no matter how hard someone tries (because the other homophones 'push' the overall letter frequency above normal levels).
So I have to admit, although other thoughts were present earlier, that - after applying a binomial tree with hypergeometric distribution values - there are only three 'candidates' of frequent letters that may fulfill the criteria to match the three double ++ formations (Z-408-frequencies):
S - 16.7 expected letters: 34.4%
L - 27.5 expected letters: 7.2%
O - 22.4 expected letters: 21.9%
The letter E, for example, had
E - 44.9 expected letters: 0.07%
To calculate the letter S, it was even necessary to set the minimum of present double letters (SS) in the cipher to an expected minimum of 6..on the other hand it was not yet possible to complete the calculation for more than three double letters ('draws'). Thus the values above are only approaches (so far I could not find any formula for hypergeomatric permutation over multiple draws..assume a bowl of different homophones drawing two symbols each time..what are the chances to draw 1, 2, 3, 4 etc. times to get at least 3 pairs of ++ ..even with such formula we still couldn't tell how many double letters are present in the cleartext).
What the observation above shows, however, is that it is very unlikely that any letter with multiple homophones is present in the cipher with 24 + symbols. Not only that those cover already 7% of the cipher but the other homophones representing the same letter have not even been considered yet. Even if the + symbol represented the letter L, there would be more homophones representing L - leading to a way too high value for the L letter frequency. Similar with O, only 22.4 letters are expected but already one of at least 3 homophones does represent 24 letters.
My conclusion to all this is that no matter how you look at it, a multiple (>2) homophone letter doesn't work out here at all. Also the letters with 2 homophones don't work as they would at least be present with 10% in the cipher (e.g. 30 letters, 24 of them a + symbol).
If we now look back to the 408, the maximum symbol there was the reversed Q symbol with 16 or 4%. Although the cipher text was shorter AND there had been fewer homophones!!
So there are two possibilities..we either can wonder about such high presence of one single homophone OOOORRRR

due to the cleartext it was absolutely inevitable to use those 24 + symbols. The only case I can think of is the case that the + symbol represents a letter which is present with ONE homophone only.
Now those would rather not be the letter Q or K etc..as they are commonly not present as multiple double letters. But there are other medium-frequent letters represented by one homophone such as C, M, P, G. Of those the letter M is expected to be one of the most frequent and indeed: The most frequent symbol of the 408 cipher was represented by nothing else but the letter M!
What I'd like to say is that it is very unlikely that any frequent letter is represented by the + symbol. This is also the reason why there is no sequence possible between the + symbols (three pairs of them occur directly after another, two with one homophone inbetween +b+). If we count the reversed Q in the 408, we get 16. In that case, however, the reversed Q or (M letter) was present with a normal frequency. If, however, such single-homophone-letter had an unexpectedly high frequency, such as 7% instead of 3%, this particular letter could be the cleartext behind +.
According to this, the letters C, M, P, G are - although above average frequency - the best candidates for the + symbol.
With double letter frequencies of
C: 0.12
G: 0.01
M: 0.05
P: 0.26
the letter P is the best double letter candidate, while the letter C is in fact closer to the overall frequency of 7%. Thus, depending on the cleartext anormalities, it is still absolutely inconclusive to tell which of those single-homophone letters shows up three times as a double letter (it could be anyone..) - and has an above average overall frequency.
HypergeomDistribution.xlsx
QT